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Jen Psaki condemned Kremlin accusations that the US was building a bioweapons lab in Ukraine and pointed out that it was Putin who had a history of using such methods to take out his enemies.
Russian defence ministry spokesman Igor Konashenkov claimed the ministry had obtained documents detailing US military-biological activities in Ukraine.
The State Centre for Research on Virology and Biotechnology in Novosibirsk Oblast, Siberia, has been a key player in Russia s bioweapon scheme for decades.