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A new paper by Hinako Komori and two academics says a specimen she found could be one of two Japanese wolves kept at Ueno Zoo in the late 19th century.

United statesCity ofUnited kingdomOscar boydHinako komoriSayaka kobayashiAlexkt martinMalcolm playfair andersonWolf canisChichibu otakiNational museum of natureUniversity of tokyoUniversity of tokyo faculty agricultureImperial household museumWakayama university faculty of educationScience department of zoology