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The facility at the Chapel Neighbourhood Centre is in addition to the one at the Glebe Centre.
A centre is also open at the Maxwell Centre in Cowdenbeath.
The centres offer testing for members of the public who have no symptoms but could be infectious. It also provides support to anyone who tests positive, with access to different financial support alongside practical measures, such as food packages.
Josie Murray, NHS Fife consultant in public health and health protection clinical lead, said: We’re basing the locations of our community testing sites on the latest data we have, allowing us to focus on targeted local areas which would benefit most from having access to testing for short periods of time.
Larbert hospital NHS Forth Valley Consultant in Public Health, Dr Jennifer Champion said: “The new variant is much more infectious and once again I urge people to follow Government advice, particularly around keeping a two metre distance, and increased hand washing. Wearing a mask does not mean that guidelines no longer apply; keeping to physical distancing is paramount in trying to stop the spread. It is so important not to let your guard down, even for a minute, to avoid becoming infected. So please remain vigilant.” The Larbert site, which has around 300 staff, has introduced additional measures to help prevent the further spread of the virus.