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Pressures were at times significant, with the nature of the forage being a key factor, researchers in Britain found.

United kingdomCity ofPam bennett skinnerSamantha hodgsonAndrea ellisPatricia harrisBryony lancasterSarah uptonNottingham trent universityRoyal dick school of veterinaryWaltham petcare science instituteSchool of animalEnvironmental science at nottingham trent universityUnequi ltdUniversity of edinburghVeterinary studies