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United kingdomJonny appleyardRoyal zoological society of scotlandWest africaRoyal zoological societyஒன்றுபட்டது கிஂக்டம்அரச ஸூலாஜிகல் சமூகம் ஆஃப் ஸ்காட்லாந்துஅரச ஸூலாஜிகல் சமூகம்