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When asked whether animals have a sense of humor, wildlife photographers Paul Joynson-Hicks and Tom Sullam respond in one word, “Completely!!!” with three exclamation points. They've seen it firsthand while crawling in the grass and hiding in the bushes. They’ve watched leopard cubs romp about the terrain, and they've seen alpha lions soften in the presence of their young. “From the smallest antelope to the oldest silver back,” says Sullam, every creature has his own kind of wit and playfulness.

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