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bioRxiv: Pierpaolo Maisano Delser, Mario Krapp, Robert Beyer, Eppie Jones, Eleanor F Miller, Anahit Hovhannisyan, Michelle Parker, Veronika Siska, Maria Teresa Vizzari, Elizabeth J Pearmain, Ivan Imaz-Rosshandler, Michela Leonardi, Gian Luigi Somma, Jason Hodgson, Eirlys Tysall, Zhe Xue, Lara Cassidy, Daniel G Bradley, Anders Eriksson, Andrea Manica This article is a preprint and has not been certified by peer review Abstract Extensive sequencing of modern and ancient human genomes has revealed that contemporary populations can be explained as the result of recent mixing of a few distinct ancestral genetic lineages. But the small number of aDNA samples that predate the Last Glacial Maximum means that the origins of these lineages are not well understood. Here, we circumvent the limited sampling by modelling explicitly the effect of climatic changes and terrain on population demography and migrations through time and space, and show that these factors are sufficient to explain the

Red seaDjibouti generalEirlys tysallMichelle parkerEppie jonesClara cassidyAnahit hovhannisyanAnders erikssonJason hodgsonGian luigi sommaPierpaolo maisano delserMario krappMichela leonardiRobert beyerEleanorf millerIvan imaz rosshandler